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Acknowledgements
The authors would like to thank to dr Beenu Jalali for her comments on the manuscript and Agnieszka Bacławska
for her technical assistance. Funded by KNOW (Leading National Research Centre) Scientific Consortium
“Healthy Animal - Safe Food”, decision of Ministry of Science and Higher Education No. 05-1/KNOW2/2015.
A.Sz-M was supported by Post-Doctoral Fellowship Program of the Japan Society for the Promotion of Science
(JSPS) (P14802).
Author contributions
A.S.z.-M. - developed the study concept, carried out experiments, analyzed and interpreted the results, and wrote
the manuscript. D.J.S., K.O. - contributed to the study concept and revised the manuscript; J.P. - participated in
isolation and culture of epithelial cells; M.S. - carried out zymography and analyzed the results.
Competing interests
The authors declare no competing interests.
Additional information
Supplementary information is available for this paper at https://doi.org/10.1038/s41598-020-58109-0.
Correspondence and requests for materials should be addressed to A.S.-M.
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